On the nomenclatural status of type genera in Coleoptera (Insecta)

doi:10.3897/zookeys.1194.106440

NIGPAS OpenIR

	On the nomenclatural status of type genera in Coleoptera (Insecta)
通讯作者	Bouchard, Patrice([email protected])
	Bouchard, Patrice 1; Bousquet, Yves 2; Davies, Anthony E.1; Cai, Chenyang2,3
	2024
发表期刊	ZOOKEYS
ISSN	1313-2989
期号	1194 页码:1-981
摘要	More than 4700 nominal family-group names (including names for fossils and ichnotaxa) are nomenclaturally available in the order Coleoptera. Since each family-group name is based on the concept of its type genus, we argue that the stability of names used for the classification of beetles depends on accurate nomenclatural data for each type genus. Following a review of taxonomic literature, with a focus on works that potentially con-tain type species designations, we provide a synthesis of nomenclatural data associated with the type genus of each nomenclaturally available family-group name in Coleoptera. For each type genus the author(s), year of publication, and page number are given as well as its current status (i.e., whether treated as valid or not) and current classification.Information about the type species of each type genus and the type species fixation (i.e., fixed originally or subsequently, and if subsequently, by whom) is also given. The orig-inal spelling of the family-group name that is based on each type genus is included, with its author(s), year, and stem. We append a list of nomenclaturally available family-group names presented in a classification scheme. Because of the importance of the Principle of Priority in zoological nomenclature, we provide information on the date of publication of the references cited in this work, when known. Several nomenclatural issues emerged during the course of this work. We therefore appeal to the community of coleopterists to submit applications to the International Commission on Zoological Nomenclature (hence-forth "Commission") in order to permanently resolve some of the problems outlined here.The following changes of authorship for type genera are implemented here (these changes do not affect the concept of each type genus): CHRYSOMELIDAE: Fulcidax Crotch, 1870 (previously credited to "Clavareau, 1913"); CICINDELIDAE: Euprosopus W.S. MacLeay, 1825 (previously credited to "Dejean, 1825"); COCCINELLIDAE: Alesia Reiche, 1848 (pre-viously credited to "Mulsant, 1850"); CURCULIONIDAE: Arachnopus Boisduval, 1835 (pre-viously credited to "Gu & eacute;rin-M & eacute;neville, 1838"); ELATERIDAE: Thylacosternus Gemminger, 1869 (previously credited to "Bonvouloir, 1871"); EUCNEMIDAE: Arrhipis Gemminger, 1869 (previously credited to "Bonvouloir, 1871"), Mesogenus Gemminger, 1869 (previously cred-ited to "Bonvouloir, 1871"); LUCANIDAE: Sinodendron Hellwig, 1791 (previously credited to "Hellwig, 1792"); PASSALIDAE: Neleides Harold, 1868 (previously credited to "Kaup, 1869"), Neleus Harold, 1868 (previously credited to "Kaup, 1869"), Pertinax Harold, 1868 (previous-ly credited to "Kaup, 1869"), Petrejus Harold, 1868 (previously credited to "Kaup, 1869"), Un-dulifer Harold, 1868 (previously credited to "Kaup, 1869"), Vatinius Harold, 1868 (previously credited to "Kaup, 1869"); PTINIDAE: Mezium Leach, 1819 (previously credited to "Curtis, 1828"); PYROCHROIDAE: Agnathus Germar, 1818 (previously credited to "Germar, 1825"); SCARABAEIDAE: Eucranium Dejean, 1833 (previously "Brull & eacute;, 1838") The following changes of type species were implemented following the discovery of older type species fixations (these changes do not pose a threat to nomenclatural sta-bility): BOLBOCERATIDAE: Bolbocerus bocchus Erichson, 1841 for Bolbelasmus Bouco-mont, 1911 (previously Bolboceras gallicum Mulsant, 1842); BUPRESTIDAE: Stigmodera guerinii Hope, 1843 for Neocuris Saunders, 1868 (previously Anthaxia fortnumi Hope, 1846), Stigmodera peroni Laporte & Gory, 1837 for Curis Laporte & Gory, 1837 (previous-ly Buprestis caloptera Boisduval, 1835); CARABIDAE: Carabus elatus Fabricius, 1801 for Molops Bonelli, 1810 (previously Carabus terricola Herbst, 1784 sensu Fabricius, 1792); CERAMBYCIDAE: Prionus palmatus Fabricius, 1792 for Macrotoma Audinet-Serville, 1832 (previously Prionus serripes Fabricius, 1781); CHRYSOMELIDAE: Donacia equise-ti Fabricius, 1798 for Haemonia Dejean, 1821 (previously Donacia zosterae Fabricius, 1801), Eumolpus ruber Latreille, 1807 for Euryope Dalman, 1824 (previously Cryptoceph-alus rubrifrons Fabricius, 1787), Galeruca affinis Paykull, 1799 for Psylliodes Latreille, 1829 (previously Chrysomela chrysocephala Linnaeus, 1758); COCCINELLIDAE: Der-mestes rufus Herbst, 1783 for Coccidula Kugelann, 1798 (previously Chrysomela scute-llata Herbst, 1783); CRYPTOPHAGIDAE: Ips caricis G.-A. Olivier, 1790 for TelmatophilusHeer, 1841 (previously Cryptophagus typhae Fall & eacute;n, 1802), Silpha evanescens Marsham, 1802 for Atomaria Stephens, 1829 (previously Dermestes nigripennis Paykull, 1798); CURCULIONIDAE: Bostrichus cinereus Herbst, 1794 for Crypturgus Erichson, 1836 (previously Bostrichus pusillus Gyllenhal, 1813); DERMESTIDAE: Dermestes trifascia-tus Fabricius, 1787 for Attagenus Latreille, 1802 (previously Dermestes pellio Linnae-us, 1758); ELATERIDAE: Elater sulcatus Fabricius, 1777 for Chalcolepidius Eschscholtz, 1829 (previously Chalcolepidius zonatus Eschscholtz, 1829); ENDOMYCHIDAE: Endo-mychus rufitarsis Chevrolat, 1835 for Epipocus Chevrolat, 1836 (previously Endomychus tibialis Gu & eacute;rin-M & eacute;neville, 1834); EROTYLIDAE: Ips humeralis Fabricius, 1787 for DacneLatreille, 1797 (previously Dermestes bipustulatus Thunberg, 1781); EUCNEMIDAE: For-nax austrocaledonicus Perroud & Montrouzier, 1865 for Mesogenus Gemminger, 1869 (previously Mesogenus mellyi Bonvouloir, 1871); GLAPHYRIDAE: Melolontha serratulaeFabricius, 1792 for Glaphyrus Latreille, 1802 (previously Scarabaeus maurus Linnaeus, 1758); HISTERIDAE: Hister striatus Forster, 1771 for Onthophilus Leach, 1817 (previ-ously Hister sulcatus Moll, 1784); LAMPYRIDAE: Ototreta fornicata E. Olivier, 1900 for Ototreta E. Olivier, 1900 (previously Ototreta weyersi E. Olivier, 1900); LUCANIDAE: Lu-canus cancroides Fabricius, 1787 for Lissotes Westwood, 1855 (previously Lissotes menalcas Westwood, 1855); MELANDRYIDAE: Nothus clavipes G.-A. Olivier, 1812 for Nothus G.-A. Olivier, 1812 (previously Nothus praeustus G.-A. Olivier, 1812); MELYRIDAE: Lagria ater Fabricius, 1787 for Enicopus Stephens, 1830 (previously Dermestes hirtusLinnaeus, 1767); NITIDULIDAE: Sphaeridium luteum Fabricius, 1787 for CychramusKugelann, 1794 (previously Strongylus quadripunctatus Herbst, 1792); OEDEMERIDAE: Helops laevis Fabricius, 1787 for Ditylus Fischer, 1817 (previously Ditylus helopioidesFischer, 1817 [sic]); PHALACRIDAE: Sphaeridium aeneum Fabricius, 1792 for OlibrusErichson, 1845 (previously Sphaeridium bicolor Fabricius, 1792); RHIPICERIDAE: Sanda-lus niger Knoch, 1801 for Sandalus Knoch, 1801 (previously Sandalus petrophya Knoch, 1801); SCARABAEIDAE: Cetonia clathrata G.-A. Olivier, 1792 for Inca Lepeletier & Audi-net-Serville, 1828 (previously Cetonia ynca Weber, 1801); Gnathocera vitticollis W. Kirby, 1825 for Gnathocera W. Kirby, 1825 (previously Gnathocera immaculata W. Kirby, 1825); Melolontha villosula Illiger, 1803 for Chasmatopterus Dejean, 1821 (previously Melolon-tha hirtula Illiger, 1803); STAPHYLINIDAE: Staphylinus politus Linnaeus, 1758 for Philon-thus Stephens, 1829 (previously Staphylinus splendens Fabricius, 1792); ZOPHERIDAE: Hispa mutica Linnaeus, 1767 for Orthocerus Latreille, 1797 (previously Tenebrio hirti-cornis DeGeer, 1775). The discovery of type species fixations that are older than those currently accepted pose a threat to nomenclatural stability (an application to the Commission is neces-sary to address each problem): CANTHARIDAE: Malthinus Latreille, 1805, MalthodesKiesenwetter, 1852; CARABIDAE: Bradycellus Erichson, 1837, Chlaenius Bonelli, 1810, Harpalus Latreille, 1802, Lebia Latreille, 1802, Pheropsophus Solier, 1834, TrechusClairville, 1806; CERAMBYCIDAE: Callichroma Latreille, 1816, Callidium Fabricius, 1775, Cerasphorus Audinet-Serville, 1834, Dorcadion Dalman, 1817, Leptura Linnaeus, 1758, Mesosa Latreille, 1829, Plectromerus Haldeman, 1847; CHRYSOMELIDAE: Amblycer-us Thunberg, 1815, Chaetocnema Stephens, 1831, Chlamys Knoch, 1801, MonomacraChevrolat, 1836, Phratora Chevrolat, 1836, Stylosomus Suffrian, 1847; COLONIDAE: Co-lon Herbst, 1797; CURCULIONIDAE: Cryphalus Erichson, 1836, Lepyrus Germar, 1817; ELATERIDAE: Adelocera Latreille, 1829, Beliophorus Eschscholtz, 1829; ENDOMYCHI-DAE: Amphisternus Germar, 1843, Dapsa Latreille, 1829; GLAPHYRIDAE: AnthypnaEschscholtz, 1818; HISTERIDAE: Hololepta Paykull, 1811, Trypanaeus Eschscholtz, 1829; LEIODIDAE: Anisotoma Panzer, 1796, Camiarus Sharp, 1878, Choleva Latreille, 1797; LYCIDAE: Calopteron Laporte, 1838, Dictyoptera Latreille, 1829; MELOIDAE: Epi-cauta Dejean, 1834; NITIDULIDAE: Strongylus Herbst, 1792; SCARABAEIDAE: AnisopliaSch & ouml;nherr, 1817, Anticheira Eschscholtz, 1818, Cyclocephala Dejean, 1821, GlycyphanaBurmeister, 1842, Omaloplia Sch & ouml;nherr, 1817, Oniticellus Dejean, 1821, Parachilia Bur-meister, 1842, Xylotrupes Hope, 1837; STAPHYLINIDAE: Batrisus Aub & eacute;, 1833, Phloe-onomus Heer, 1840, Silpha Linnaeus, 1758; TENEBRIONIDAE: Bolitophagus Illiger, 1798, Mycetochara Gu & eacute;rin-M & eacute;neville, 1827.Type species are fixed for the following nominal genera: ANTHRIBIDAE: Decata-phanes gracilis Labram & Imhoff, 1840 for Decataphanes Labram & Imhoff, 1840; CARA-BIDAE: Feronia erratica Dejean, 1828 for Loxandrus J.L. LeConte, 1853; CERAMBYCIDAE: Tmesisternus oblongus Boisduval, 1835 for Icthyosoma Boisduval, 1835; CHRYSOMEL-IDAE: Brachydactyla annulipes Pic, 1913 for Pseudocrioceris Pic, 1916, Cassida viridisLinnaeus, 1758 for Evaspistes Gistel, 1856, Ocnoscelis cyanoptera Erichson, 1847 for Oc-noscelis Erichson, 1847, Promecotheca petelii Gu & eacute;rin-M & eacute;neville, 1840 for PromecothecaGu & eacute;rin- M & eacute;neville, 1840; CLERIDAE: Attelabus mollis Linnaeus, 1758 for DendroplanetesGistel, 1856; CORYLOPHIDAE: Corylophus marginicollis J.L. LeConte, 1852 for Corylo-phodes A. Matthews, 1885; CURCULIONIDAE: Hoplorhinus melanocephalus Chevrolat, 1878 for Hoplorhinus Chevrolat, 1878; Sonnetius binarius Casey, 1922 for Sonnetius Ca-sey, 1922; ELATERIDAE: Pyrophorus melanoxanthus Cand & egrave;ze, 1865 for Alampes Cham-pion, 1896; PHYCOSECIDAE: Phycosecis litoralis Pascoe, 1875 for Phycosecis Pascoe, 1875; PTILODACTYLIDAE: Aploglossa sallei Gu & eacute;rin-M & eacute;neville, 1849 for AploglossaGu & eacute;rin-M & eacute;neville, 1849, Colobodera ovata Klug, 1837 for Colobodera Klug, 1837; PTIN-IDAE: Dryophilus anobioides Chevrolat, 1832 for Dryobia Gistel, 1856; SCARABAEIDAE: Achloa helvola Erichson, 1840 for Achloa Erichson, 1840, Camenta obesa Burmeister, 1855 for Camenta Erichson, 1847, Pinotus talaus Erichson, 1847 for Pinotus Erichson, 1847, Psilonychus ecklonii Burmeister, 1855 for Psilonychus Burmeister, 1855.New replacement name: CERAMBYCIDAE: Basorus Bouchard & Bousquet, nom. nov.for Sobarus Harold, 1879.New status: CARABIDAE: KRYZHANOVSKIANINI Deuve, 2020, stat. nov. is giv-en the rank of tribe instead of subfamily since our classification uses the rank of subfamily for PAUSSINAE rather than family rank; CERAMBYCIDAE: Amymoma Pas-coe, 1866, stat. nov. is used as valid over Neoamymoma Marinoni, 1977, HolopterusBlanchard, 1851, stat. nov. is used as valid over Proholopterus Monn & eacute;, 2012; CUR-CULIONIDAE: Phytophilus Sch & ouml;nherr, 1835, stat. nov. is used as valid over the un-necessary new replacement name Synophthalmus Lacordaire, 1863; EUCNEMIDAE: Nematodinus Lea, 1919, stat. nov. is used as valid instead of Arrhipis Gemminger, 1869, which is a junior homonym.Details regarding additional nomenclatural issues that still need to be resolved are included in the entry for each of these type genera: BOSTRICHIDAE: Lyctus Fabricius, 1792; BRENTIDAE: Trachelizus Dejean, 1834; BUPRESTIDAE: PristipteraDejean, 1833; CANTHARIDAE: Chauliognathus Hentz, 1830, Telephorus Sch & auml;ffer, 1766; CARABIDAE: Calathus Bonelli, 1810, Cosnania Dejean, 1821, Dicrochile Gu & eacute;rin-M & eacute;neville, 1847, Epactius D.H. Schneider, 1791, Merismoderus Westwood, 1847, Polyhirma Chaudoir, 1850, Solenogenys Westwood, 1860, Zabrus Clairville, 1806; CE-RAMBYCIDAE: Ancita J. Thomson, 1864, Compsocerus Audinet-Serville, 1834, Dorca-dodium Gistel, 1856, Glenea Newman, 1842; Hesperophanes Dejean, 1835, NeoclytusJ. Thomson, 1860, Phymasterna Laporte, 1840, Tetrops Stephens, 1829, ZygoceraErichson, 1842; CHRYSOMELIDAE: Acanthoscelides Schilsky, 1905, Corynodes Hope, 1841, Edusella Chapuis, 1874; Hemisphaerota Chevrolat, 1836; Physonota Boheman, 1854, Porphyraspis Hope, 1841; CLERIDAE: Dermestoides Sch & auml;ffer, 1777; COCCI-NELLIDAE: Hippodamia Chevrolat, 1836, Myzia Mulsant, 1846, Platynaspis L. Redten-bacher, 1843; CURCULIONIDAE: Coeliodes Sch & ouml;nherr, 1837, Cryptoderma Ritsema, 1885, Deporaus Leach, 1819, Epistrophus Kirsch, 1869, Geonemus Sch & ouml;nherr, 1833, Hylastes Erichson, 1836; DYTISCIDAE: Deronectes Sharp, 1882, Platynectes R & eacute;gim-bart, 1879; EUCNEMIDAE: Dirhagus Latreille, 1834; HYBOSORIDAE: Ceratocanthus A. White, 1842; HYDROPHILIDAE: Cyclonotum Erichson, 1837; LAMPYRIDAE: LuciolaLaporte, 1833; LEIODIDAE: Ptomaphagus Hellwig, 1795; LUCANIDAE: LeptinopterusHope, 1838; LYCIDAE: Cladophorus Gu & eacute;rin-M & eacute;neville, 1830, Mimolibnetis Kazant-sev, 2000; MELOIDAE: Mylabris Fabricius, 1775; NITIDULIDAE: Meligethes Stephens, 1829; PTILODACTYLIDAE: Daemon Laporte, 1838; SCARABAEIDAE: Allidiostoma Ar-row, 1940, Heterochelus Burmeister, 1844, Liatongus Reitter, 1892, Lomaptera Gory & Percheron, 1833, Megaceras Hope, 1837, Stenotarsia Burmeister, 1842; STAPHYLIN-IDAE: Actocharis Fauvel, 1871, Aleochara Gravenhorst, 1802; STENOTRACHELIDAE: Stenotrachelus Berthold, 1827; TENEBRIONIDAE: Cryptochile Latreille, 1828, He-liopates Dejean, 1834, Helops Fabricius, 1775.First Reviser actions deciding the correct original spelling: CARABIDAE: Aristoch-roodes Marcilhac, 1993 (not Aritochroodes); CERAMBYCIDAE: Dorcadodium Gistel, 1856 (not Dorcadodion), EVODININI Zamoroka, 2022 (not EVODINIINI); CHRYSOMELI-DAE: Caryopemon Jekel, 1855 (not Carpopemon), Decarthrocera Laboissi & egrave;re, 1937 (not Decarthrocerina); CICINDELIDAE: Odontocheila Laporte, 1834 (not Odontacheila); CLERI-DAE: CORMODINA Bartlett, 2021 (not CORMODIINA), Orthopleura Spinola, 1845 (not Or-thoplevra, not Orthopleuva); CURCULIONIDAE: Arachnobas Boisduval, 1835 (not Arach-nopus), Palaeocryptorhynchus Poinar, 2009 (not Palaeocryptorhynus); DYTISCIDAE: Ambarticus Yang et al., 2019 and AMBARTICINI Yang et al. , 2019 (not Ambraticus, not AMBRATICINI); LAMPYRIDAE: Megalophthalmus G.R. Gray, 1831 (not Megolophthalmus, not Megalopthalmus); SCARABAEIDAE: Mentophilus Laporte, 1840 (not Mintophilus, not Minthophilus), Pseudadoretus dilutellus Semenov, 1889 (not P. ditutellus).While the correct identification of the type species is assumed, in some cases ev-idence suggests that species were misidentified when they were fixed as the type of a particular nominal genus. Following the requirements of Article 70.3.2 of the Inter-national Code of Zoological Nomenclature we hereby fix the following type species (which in each case is the taxonomic species actually involved in the misidentifica-tion): ATTELABIDAE: Rhynchites cavifrons Gyllenhal, 1833 for Lasiorhynchites Jekel, 1860; BOSTRICHIDAE: Ligniperda terebrans Pallas, 1772 for Apate Fabricius, 1775; BRENTIDAE: Ceocephalus appendiculatus Boheman, 1833 for Uroptera Berthold, 1827; BUPRESTIDAE: Buprestis undecimmaculata Herbst, 1784 for Ptosima Dejean, 1833; CA-RABIDAE: Amara lunicollis Schi & oslash;dte, 1837 for Amara Bonelli, 1810, Buprestis connexusGeoffroy, 1785 for Polistichus Bonelli, 1810, Carabus atrorufus Str & oslash;m, 1768 for PatrobusDejean, 1821, Carabus gigas Creutzer, 1799 for Procerus Dejean, 1821, Carabus teuto-nus Schrank, 1781 for Stenolophus Dejean, 1821, Carenum bonellii Westwood, 1842 for Carenum Bonelli, 1813, Scarites picipes G.-A. Olivier, 1795 for Acinopus Dejean, 1821, Trigonotoma indica Brull & eacute;, 1834 for Trigonotoma Dejean, 1828; CERAMBYCIDAE: Cer-ambyx lusitanus Linnaeus, 1767 for Exocentrus Dejean, 1835, Clytus supernotatus Say, 1824 for Psenocerus J.L. LeConte, 1852; CICINDELIDAE: Ctenostoma jekelii Chevrolat, 1858 for Ctenostoma Klug, 1821; CURCULIONIDAE: Cnemogonus lecontei Dietz, 1896 for Cnemogonus J.L. LeConte, 1876; Phloeophagus turbatus Sch & ouml;nherr, 1845 for Phloeoph-agus Sch & ouml;nherr, 1838; GEOTRUPIDAE: Lucanus apterus Laxmann, 1770 for Lethrus Sco-poli, 1777; HISTERIDAE: Hister rugiceps Duftschmid, 1805 for Hypocaccus C.G. Thom-son, 1867; HYBOSORIDAE: Hybosorus illigeri Reiche, 1853 for Hybosorus W.S. MacLeay, 1819; HYDROPHILIDAE: Hydrophilus melanocephalus G.-A. Olivier, 1793 for EnochrusC.G. Thomson, 1859; MYCETAEIDAE: Dermestes subterraneus Fabricius, 1801 for Myce-taea Stephens, 1829; SCARABAEIDAE: Aulacium carinatum Reiche, 1841 for Mentoph-ilus Laporte, 1840, Phanaeus vindex W.S. MacLeay, 1819 for Phanaeus W.S. MacLeay, 1819, Ptinus germanus Linnaeus, 1767 for Rhyssemus Mulsant, 1842, Scarabaeus lati-pes Gu & eacute;rin-M & eacute;neville, 1838 for Cheiroplatys Hope, 1837; STAPHYLINIDAE: Scydmaenus tarsatus P.W.J. M & uuml;ller & Kunze, 1822 for Scydmaenus Latreille, 1802.New synonyms: CERAMBYCIDAE: CARILIINI Zamoroka, 2022, syn. nov. of ACMAEO-PINI Della Beffa, 1915, DOLOCERINI & Ouml;zdikmen, 2016, syn. nov. of BRACHYPTEROMINI Sama, 2008, PELOSSINI Tavakilian, 2013, syn. nov. of LYGRINI Sama, 2008, PROHOL-OPTERINI Monn & eacute;, 2012, syn. nov. of HOLOPTERINI Lacordaire, 1868.
关键词	Beetles catalogue classification family-group name nomenclature publication dates type genus type species
DOI	10.3897/zookeys.1194.106440
收录类别	SCI
语种	英语
关键词[WOS]	GENUS-GROUP NAMES ; EARLY CRETACEOUS WEEVILS ; FAMILY-GROUP NAMES ; TRIB.-NOV. COLEOPTERA ; NET-WINGED BEETLES ; SUPERFAMILY CURCULIONOIDEA COLEOPTERA ; BALTIC AMBER ; MOLECULAR PHYLOGENY ; ANNOTATED KEY ; CORYLOPHIDAE COLEOPTERA
WOS研究方向	Zoology
WOS类目	Zoology
WOS记录号	WOS:001333688300001
出版者	PENSOFT PUBLISHERS
文献类型	期刊论文
条目标识符	http://ir.nigpas.ac.cn/handle/332004/44501
专题	中国科学院南京地质古生物研究所
通讯作者	Bouchard, Patrice
作者单位	1.Canadian Natl Collect Insects Arachnids & Nematode, Agr & Agri Food Canada, 960 Carling Ave, Ottawa, ON K1A 0C6, Canada 2.Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, Nanjing 210008, Peoples R China 3.Chinese Acad Sci, Ctr Excellence Life & Paleoenvironm, Nanjing 210008, Peoples R China
推荐引用方式 GB/T 7714	Bouchard, Patrice,Bousquet, Yves,Davies, Anthony E.,et al. On the nomenclatural status of type genera in Coleoptera (Insecta)[J]. ZOOKEYS,2024(1194):1-981.
APA	Bouchard, Patrice,Bousquet, Yves,Davies, Anthony E.,&Cai, Chenyang.(2024).On the nomenclatural status of type genera in Coleoptera (Insecta).ZOOKEYS(1194),1-981.
MLA	Bouchard, Patrice,et al."On the nomenclatural status of type genera in Coleoptera (Insecta)".ZOOKEYS .1194(2024):1-981.